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Graphic showing how pilots determine an enemy plane location relative to their own planes

As can be seen in the graphic above a very simple yet effective method
of locating enemy planes in the sky was developed and used by allied
aircrews. This system became a workable solution to the old method
of hand signals and wing wig-wags when the use of inter-plane radio
communication became a widespread, although not completely reliable,
part of all allied aircraft. While the Allies were perfecting this type of
air-to-air and air-to-ground communication the Japanese were not as
quick to see the advantage. They not only retained the hand signals but
due to a lack of reliable radios were actually known to remove their
radios in order to lighten their airplanes. A mistake that proved fatal.

The best position to attack a plane was from 6 o'clock astern with a
deflection no greater than 20 degrees. What does "a deflection no
greater than 20 degrees" mean? If you look at the graphic above and
assume that 6 o'clock astern equals zero degrees then a 20 degree
deflection would be 20 degrees from that point or from 5 to 7 o'clock.
The same applied to a frontal attack (11 to 1 o'clock) but since the
aircraft were approaching each other at high speed the optimal time to
fire was very short.

Once the angle of deflection went beyond 20 degrees a number of
factors came into play and the pilot had to judge his speed, the speed
and direction of the enemy and make appropriate adjustments to his
angle of attack before firing. For instance if he were attacking from
the 9 o'clock position it would be necessary to "lead" the enemy
aircraft in order to hit it. In other words he wouldn't shoot directly at
the airplane because by the time the bullets got there the airplane
would have flown ahead of that point. He would fire at a point in front
of the enemy airplane so that by the time the bullets got there the
enemy plane would also have reached that point. Combine this with
factors such as whether he is above or below the enemy airplane,
approaching from in front or behind and whether the enemy is taking
evasive action and you begin to understand some of the difficulties in
shooting down a warplane.

Diagram of Cone of Fire

Another factor that came into play was something called the "cone
of fire". All fighters with machine guns in the wings had those guns
aligned so that when they fired all the trajectories would converge at
a point about 200-300 yards away. At that point would be the highest
concentration of bullets. Beyond that the bullets started to diverge and
lose effectiveness. In addition since the wings were mounted below the
center-line of the airplane it was necessary to "pull the nose through
the enemy plane" when firing thereby losing sight of the target as
he fired. He then had to drop the nose in order to reaquire the target
and see if any damage had been done. The one plane that was different
was the P-38 Lightning. All the machine guns were mounted close
together in the nose and therefore its cone of fire was fully concentrated
all the way out. Taking all this into consideration and you can see that
it was incredibly difficult to shoot down another warplane. That's what
makes the accomplishments of the aces so remarkable.

Diagram explaining what the angle of deflection is

Thin grey divider line

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Small early AF insignia